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4. Results and Discussion

4.1. Faunal spectrum

Figure 2. Identified Marine invertebrates at Clos des Châtaigniers (Image credit: C. Mougne)
1: Mytilus edulis (L = 35 mm), 2: Acanthocardia tuberculata (L = 51 mm), 3: Cerastoderma edule (L = 27 mm), 4: Solen marginatus (L = 16 mm), 5: Scrobicularia plana (L = 16 mm), 6: Ruditapes decussatus (L = 4 mm), 7: Barnea candida (L = 13 mm), 8: Macoma balthica (L = 18 mm), 9: Donax vittatus (L = 18 mm), 10: Mactra sp. (L = 18 mm), 11: Spisula solida (L = 33 mm), 12: Spisula subtruncata (L = 14 mm), 13: Ostrea edulis (L = 25 mm), 14: Aequipecten opercularis (L = 8 mm), 15: Mimaclamys varia (L = 10 mm), 16: Corbula gibba (L = 9 mm), 17: Phaxas pellucidus (L = 9 mm), 18: Anomia ephippium (L = 15 mm ), 19: Abra sp. (L = 4 mm), 20: Gari sp. (L = 6 mm), 21: Gibbula umbilicalis (L = 13 mm), 22: Littorina littorea (L = 22 mm), 23: Buccinum undatum (L = 32 mm), 24: Nucella lapillus (L = 22 mm), 25: Nassarius reticulatus (L = 22 mm), 26: Ocenebra erinaceus (L = 34 mm), 27: Bela powisiana (L = 10 mm), 28: Epitonium clathrus (L = 10 mm), 29: Lacuna pallidula (L = 7 mm), 30: Littorina obtusata (L = 7 mm), 31: Test of urchin (L = 6 mm) 32: Balanus sp. (L = 11 mm). [Static image]

Thirty species of marine shells (20 bivalves, 10 gastropods), one sea urchin and one crustacean were identified (Figure 2) at the site of Clos des Châtaigniers. Mussels (Mytilus edulis) represent the majority of marine shell remains in all three types of quantification used (98% of the NISP, 75% of the MNI and 94% of total weight) (Figure 3). The high percentage of NISP for this species is linked to high levels of fragmentation. As a consequence, the relation between the Number of Identified Specimens and the Minimum Number of Individuals (221752/1490) shows that, for a single mussel individual, 149 remains were counted. It is thus often difficult to obtain total lengths and to describe the collecting strategies. From these correlations between total length and a small part near the teeth, 214 total lengths (RV = 105; LV = 109) have been reconstructed. These measurements ranged between 27 and 65mm, with a mean of 43.5mm, corresponding to adult individuals (Figure 4).

Figure 3: Spectrum of the marine invertebrates at Clos des Châtaigniers
Latin name Common name NISP MNI Weight (g)
Bivalves
Mytilus edulis common mussel 221752 1490 6188.24
Ostrea edulis flat oyster 33 4 5.81
Anomia ephippium saddle oyster 1 1 0.18
Aequipecten opercularis queen scallop 1 1 0.01
Mimachlamys varia fan scallop 3 1 0.08
Solen marginatus grooved razor shell 10 4 1.96
Phaxas pellucidus   7 5 0.04
Acanthocardia tuberculata   2 2 19.19
Acanthocardia sp.   115 3 16.48
Cerastoderma edule common cockle 107 16 12.65
Cerastoderma sp. cockle 186 8 110.47
Barnea candida   669 36 11.8
Donax vittatus beam clam 1 1 0.36
Donax sp. beam clam 275 25 4.64
Gari sp.   10 3 0.11
Macoma balthica   86 33 5.05
Mactra sp.   29 11 0.88
Spisula solida Atlantic surf clam 1 1 2.21
Spisula subtruncata   1 1 0.03
Spisula sp.   131 54 1.92
Scrobicularia plana peppery furrow shell 149 4 4.09
Abra sp.   39 25 0.24
Corbula gibba   22 8 0.68
Ruditapes decussatus European carpet clam 1 1 0.08
Cardiidae   22   0.71
Mactridae   891 7 5.83
Pectinidae   2   0.03
Semelidae   278   3.21
Tellinidae   188   1.8
Veneroida   549   4.51
Gastropods
Littorina littorea common periwinkle 336 123 123.78
Littorina obtusata flat periwinkle 18 11 1.53
Littorina sp.   44 41 0.55
Lacuna pallidula pale lacuna 1 1 0.12
Gibbula umbilicalis flat top shell 16 13 6.54
Gibbula sp. top shell 48 3 1.77
Buccinum undatum common whelk 4 2 4.77
Nassarius reticulatus netted dog whelk 27 5 5.21
Nassarius sp.   5 2 0.19
Nucella lapillus dogwhelk 57 20 39.01
Ocenebra erinaceus sting winkle 28 6 6.39
Bela powisiana   1 1 0.04
Epitonium clathrus common wentletrap 1 1 0.1
Unspecified gastropods   19   1.12
Unspectified molluscs   15   0.22
Marine Molluscs total   226181 1974 6594.63
Balanus sp. barnacle 35566 5927 187.39
Crustacean total   35566  5927 187.39
Unspecified urchin urchin 2 1 0.03
Echinoderm total   2  1 0.03

Figure 4
Figure 4: Distribution of reconstructed length (1) classes (mm) of Mytilus edulis, from a small part near the teeth (2). (N = number of valves) (Image credit (mussel): L. Quesnel)

Seven other identified marine shells are edible and may have been part of the diet of the inhabitants of Clos des Châtaigniers: Littorina littorea, Barnea candida, Macoma balthica, Donax sp., Abra sp., Spisula sp. and Cerastoderma sp. (Figure 3). Nonetheless, the shells of these bivalves measured less than 20mm, a size typically rejected as being too small for human consumption. Thus the presence of these six species does not appear to be linked to human diet. The dogwhelk (Nucella lapillus) is represented by 20 medium-sized individuals (between 20 and 40mm). Half of them, however, show marks of marine worms of Polydora-type in their internal shell. These taphonomic modifications show that specimens of dogwhelk arrived dead and empty, without flesh, to the site. Twenty-one other species (Anomia ephippium, Ostrea edulis, Solen marginatus, Acanthocardia tuberculata, Scrobicularia plana, Spisula solida, Buccinum undatum, Aequipecten opercularis, Mimachlamys varia, Ruditapes decussatus, Gari sp., Phaxas pellucidus, Lacuna pallidula, Littorina obtusata, Bela powisiana, Nassarius reticulatus, Epitonium clathrus, Corbula gibba, Ocenebra erinaceus, Mactra sp. and Gibbula sp.) are represented by only 1 to 13 individuals (Figure 3). The small quantities of these marine invertebrates are unlikely to indicate intentional collecting for human consumption. In addition, some of these species are small in size and evidence shows that deposited specimens were already dead when they arrived at the site. Only one fragment of sea urchin, species unidentified, was found (Figure 2, 31). Finally, some barnacles, belonging to the order of Cirripedia crustaceans, were observed (Figure 2, 32) and probably brought to the site during the transport of the mussels.

In summary, the size and quantity of mussel remains show that this shellfish was consumed by past inhabitants. However, the consumption of the other thirty-one identified marine invertebrates cannot be proved because of their presence in small quantities (Figure 3), small sizes and/or observed taphonomic modifications.